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A QUANTITATIVE ANALYSIS OF POLLEN VARIATION IN TWO SOUTHERN CALIFORNIA PERENNIAL HELIANTHUS (HELIANTHEAE: ASTERACEAE)
J. Mark Porter and Naomi Fraga
Rancho Santa Ana Botanic Garden
1500 N. College Ave., Claremont, CA 91711

ABSTRACT: A quantitative analysis of pollen from Helianthus californicus, H. nuttallii var. nuttallii, H. nuttallii var. parishii, and H. "Newhall Ranch" is undertaken. Equatorial diameter, polar diameter, length and width of colpi, pore diameter, spine length, number of micropores at the base of spines, and number of spines in a direct line along the equator of the grain between apertures were measured for 34 collections of these taxa. Multivariate analysis of variance (MANOVA) of continuous variables reveals a significant difference among the taxa. Post hoc multiple pairwise comparisons show that the difference discovered using MANOVA is due largely to the differences between H. californicus and H. nuttallii (subsp. nuttallii and parishii do not differ). The Newhall Ranch population differed significantly in some traits with H. californicus; however, differed in others with H. nuttallii. This intermediate relationship is consistent with chromosome counts, which are also intermediate to the counts available for H. californicus and H. nuttallii. While not conclusive, these data are consistent with the hypothesis that the Newhall Ranch population is neither H. californicus nor H. nuttallii, but perhaps an intermediate polyploid, linking the two.

KEYWORDS: Helianthus californicus, Helianthus nuttallii subsp. nuttallii, Helianthus nuttallii subsp. parishii, palynology, quantitative analysis, rare taxa.

THE CARBONATE LICHEN FLORA OF CACTUS FLATS IN THE SAN BERNARDINO MOUNTAINS, SAN BERNARDINO COUNTY, CALIFORNIA, USA
Kerry Knudsen
The Herbarium, Department of Botany & Plant Sciences
University of California, Riverside, CA, 92521-0124

ABSTRACT: Thirty nine species of lichen in twenty-four genera are reported from carbonate rock and soil in the Cactus Flats area of the San Bernardino Mountains. Lichenochora xanthoriae Triebel & Rambold, Polysporina uceolata (Anzi) Brodo and Verrucaria zamenhofiana Clauzade & Cl. Roux are reported as new to California. A species new to science discovered at Cactus Flats, Verrucaria bernardinensis Breuss in ed., is discussed in anticipation of its publication.

KEYWORDS: Cactus Flats, calciphiles, Carbonate Habitat Management Strategy, carbonate lichen flora, carbonate vascular flora, lichens, lichenized ascomycetes, lichenicoles, limestone, Mojave desert, San Bernardino National Forest, San Bernardino Mountains

A BIBLIOGRAPHY OF FLORISTICS IN SOUTHERN CALIFORNIA: ADDENDUM NUMBER TWO
Robert F. Thorne
Rancho Santa Ana Botanic Garden
1500 N. College Ave., Claremont, CA 91711

ABSTRACT: A bibliography of floristics in southern California was published in Crossosoma Volume 24, Numbers l and 2, in 1999 (for Spring-Summer and Autumn-Winter 1998). At that time it was realized that some important references might have been overlooked and that subsequently many more pertinent to floristics in southern California would be published each year. It was hoped that at least once a year an addendum could be published in Crossosoma to bring the bibliography up to date. An addendum was published in Volume 27, Number 2, in 2002 (for Fall-Winter 2001). The following additional references have been gleaned since that time from the library and literature or as suggested by members of Southern California Botanists. As in the previous publications, they are divided into Part 1 for the Entire region and Part 2 for Literature pertinent to local areas.